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In the same study, seeds of Tropeaolum hjertingii Sparre and Tropaeolum minus L. were shown to produce only benzyl ITC upon myrosinase hydrolysis, confirming the presence of only benzyl GL (3) in the seeds [59]. Finally, it has been established that benzyl (3), 4-methoxybenzyl (5), 1-methylethyl (26), 1-methylpropyl (27) and 2-methylpropyl (28) GLs are present in the seeds of Tropaeolum cochabambae Buch [59]. 2. GLUCOSINOLATE DISTRIBUTION IN THE ORDERS MALPIGHIALES, CELASTRALES, CARYOPHYLLALES, AND GENTIANALES The occurrence of GLs in some of the following plant orders are questionable and would need further confirmations.

1970). 2-Hydroxy-2-methylpropyl glucosinolate in Reseda alba. Phytochemistry 9, 599-600. , Bennett, R. , Botting, N. , Davidson, N. , Robertson, A. A. , Wallsgrove, R. , (2001). High-performance liquid chromatographic separation of natural and synthetic desulfoglucosinolates and their chemical validation by UV, NMR and chemical ionisation-MS methods. Phytochem. Anal. 12, 226-242. , (1979). Isolation of glucosinolates and the identification of O(α-L-rhamnopyranosyloxy)benzylglucosinolate from Reseda odorata.

However, these studies also typically conclude that individual cultivars respond differently to 40 Mark W. Farnham and Sandra E. Branham varying sulfur fertility [29], and more than one study has found that the effects of sulfur fertilization can be largely influenced by nitrogen nutrition [31, 32]. Although increasing sulfur fertilization sometimes increases glucosinolate concentration in broccoli heads, applications of selenium tend to decrease glucosinolate concentrations [19, 33]. This is believed to occur due to the fact that selenium (taken up as SeO4: selenate) replaces sulfur and interferes with normal glucosinolate metabolism [34].

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