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Synapsis being the union of homologues would result in the formation of the bivalent chromosomes Aa, Bb, Cc, Dd, which would again be resolved into their components by the reducing division. Each of the ripe germ-cells arising from the reduction divisions must receive one member from each mosomes A, and B, C, a, b, c, d, of the synaptic pairs, but there are sixteen possible combinations of C, d; a, b, C, D; a, B, c, D; a, B, C, d; a, b, c, d; and their conjugates A, b, c, d; c, a, c, d; B, Hence d; a, b, C, d; a, b, c, D; A, B, C, d; A, b, c, D; A, B, C, D.

On the contrary, the germ-cells are equivalent, as all shown by tions the fact that all combina- produce similar offspring which in turn are similar to the parent. suggestion made by Bateson in The an- morph were alone produced by the male and the other by the female we should have a species consisting only of heterozygotes," which would come true as long as bred together, at first sight seems logically applicable to For such an idea, however, can find no cytological justification, since if any reduction occurs both chromosomes occur in both male and female germ-cells in equal numbers; and further, the evidence is in favor of a great variety of combinations of maternal and paternal chrothese cases.

6 de "Intracellulare Vries, Jena, 1889. necessary principle in truly intensive research in the study of heredity, then its highest purpose would be achieved. Later publications will attempt to illuminate the activity of lines which vary polymodally. I have investigated only unimodal variation in this paper,'^ in order to present my concept in its simplest instance. The lies Descen- Pangenesis," thought which under- train of this investigation is simplicity most clearly its cited unvery concept changed.

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